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Original Article

Identity of Spirometra theileri from a Leopard (Panthera pardus) and Spotted Hyena (Crocuta crocuta) in Tanzania

Keeseon S. Eom1, Hansol Park1, Dongmin Lee1, Seongjun Choe1, Yeseul Kang1, Mohammed Mebarek Bia1, Barakaeli Abdieli Ndosi1,4, Tilak Chandra Nath1,5, Chatanun Eamudomkarn2, Julius Keyyu3, Robert Fyumagwa3, Simon Mduma3, Hyeong-Kyu Jeon1,*
The Korean Journal of Parasitology 2019;57(6):639-645.
Published online: December 31, 2019

1Department of Parasitology, Parasitology Research Center and Parasite Resource Bank, Chungbuk National University School of Medicine, Cheongju 28644, Korea

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*Corresponding author (jeonhk@chungbuk.ac.kr)
• Received: September 15, 2019   • Revised: October 21, 2019   • Accepted: October 21, 2019

Copyright © 2019 by The Korean Society for Parasitology and Tropical Medicine

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Identity of Spirometra theileri from a Leopard (Panthera pardus) and Spotted Hyena (Crocuta crocuta) in Tanzania
Korean J Parasitol. 2019;57(6):639-645.   Published online December 31, 2019
DOI: https://doi.org/10.3347/kjp.2019.57.6.639
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Identity of Spirometra theileri from a Leopard (Panthera pardus) and Spotted Hyena (Crocuta crocuta) in Tanzania
Korean J Parasitol. 2019;57(6):639-645.   Published online December 31, 2019
DOI: https://doi.org/10.3347/kjp.2019.57.6.639
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Identity of Spirometra theileri from a Leopard (Panthera pardus) and Spotted Hyena (Crocuta crocuta) in Tanzania
Image Image
Fig. 1 Gravid proglottids of Spirometra theileri (Tanzania origin) collected from a leopard (Panthera pardus) (A) and spotted hyena (Crocuta crocuta) (B). (A, B) Whole mounted specimens of proglottids showing the genital pore (GP), vaginal pore (VP), and uterus (U) (×40), (C) Longitudinal section of a gravid proglottid showing the cirrus sac (CS), seminal vesicle (SV), and uterus (U) (H&E stain).
Fig. 2 Phylogenetic relationship of Spirometra species based on cox1 and nad1 sequences. Numbers on branches represent bootstrap values for maximum likelihood (ML) and the support value of Bayesian inference (BI). GenBank No. MK955901 and MN244299 are registered as S. theileri by complete mitochondrial cox1 and nad1 sequences.

Fig. 1

Fig. 2

Identity of Spirometra theileri from a Leopard (Panthera pardus) and Spotted Hyena (Crocuta crocuta) in Tanzania

Morphological features of Spirometra theileri and Spirometra sp.

Organs Morphological features S. theireli1 Spirometra sp.2 Spirometra sp.3 Others
Size (mm)
Scolex Spatulate 0.25–0.3 - - Diameter
0.53–0.86 - - Length
Proglottids4 (n=10) Trapezoid 0.4–2.4 0.60–0.65 0.4–1.0 Width
0.02–1.9 0.15–0.25 0.15–0.25 Length
Uterus (n=10) Coiling 3–4.5 3–4.5 3–4.5 Loops
Eggs (n=10) Operculate 0.057 0.054–0.059 0.054–0.059 Length
0.034 0.033–0.035 0.033–0.035 Width
Cirrus pouch (n=10) Oval 0.16 0.16 0.16 Length
0.13 0.13 0.13 Width
Cirrus (n=10) Cylinderical 0.3 0.3 0.3 Length
0.13 0.13 0.13 Diameter
Seminal vesicle (n=10) Elliptical - 0.13 0.13 Length
0.14 0.14 Width
Vaginal opening (n=10) Crescentic - 0.25 0.25 Width
Testes (n=10) Spherical 0.11–0.15:0.06–0.08 0.11–0.12:0.04–0.06 0.11–0.12:0.04–0.06 Diameter

1Baer (1926).

2This study (Tanzania origin collected from a leopard).

3This study (Tanzania origin collected from a spotted hyena).

4Range from the first segment to the end of the segment of the strobila.

Pairwise sequence differences of mitochondrial cox1 and nad1 genes of Spirometra species of Tanzania origin and various Spirometra species, Dibothriocephalus latus and D. nihonkaiense

−1 −2 −3 −4 −5 −6
(1) Spirometra species (Tanzania)*
(2) S. decipiens (KJ599679) 9.6/13.0
(3) S. ranarum (MH298843/MH298844) 9.8/12.6 2.1/2.1
(4) S. erinaceieuropaei (KJ599680) 10.1/12.0 9.4/8.8 8.8/10.0
(5) Sparganum proliferum (AB015753) 12.2/− 12.1/− 12.2/− 12.6/−
(6) D. latus (DQ985706) 16.7/20.3 17.4/22.0 17.4/21.7 16.9/21.7 16.9/−
(7) D. nihonkaiense (EF420138) 17.8/21.8 17.5/22.3 17.8/22.0 17.6/21.7 17.6/7.2 7.2/7.2

*S. theileri (GenBank No. MK955901 for cox1 and MN244299 for nad1).

Genetic diversity of mitochondrial DNA sequences of Spirometra species including S. erinaceieuropaei, S. decipiens, S. ranarum and S. theileri

V-S C-S Sin-S Parsi-S Syn-S NonSyn-S Sq-D Hd±S.D. Pi±S.D.
cox1 (1,566 bp) 227 1,339 169 59 374.75 1188.25 0–0.0101 1.000±0.177 0.08280±0.01946
nad1 (891 bp) 157 734 116 41 209.00 682.00 0–0.0130 1.000±0.177 0.09914±0.02443

V-S: variable site, C-S: conserved site, Sin-S: singleton site, Parsi-S: parsimony informative site, Syn-S: synonymous site, NonSyn-S: nonsynonymous site, Sq-D: sequence divergence, Hd: haplotype diversity, Pi: nucleotide diversity.

Table 1 Morphological features of Spirometra theileri and Spirometra sp.

Baer (1926).

This study (Tanzania origin collected from a leopard).

This study (Tanzania origin collected from a spotted hyena).

Range from the first segment to the end of the segment of the strobila.

Table 2 Pairwise sequence differences of mitochondrial cox1 and nad1 genes of Spirometra species of Tanzania origin and various Spirometra species, Dibothriocephalus latus and D. nihonkaiense

S. theileri (GenBank No. MK955901 for cox1 and MN244299 for nad1).

Table 3 Genetic diversity of mitochondrial DNA sequences of Spirometra species including S. erinaceieuropaei, S. decipiens, S. ranarum and S. theileri

V-S: variable site, C-S: conserved site, Sin-S: singleton site, Parsi-S: parsimony informative site, Syn-S: synonymous site, NonSyn-S: nonsynonymous site, Sq-D: sequence divergence, Hd: haplotype diversity, Pi: nucleotide diversity.

Table 1

Table 2

Table 3

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